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1955-1961 - B.Sc. (Hons) in Botany at University College London
1961-1962 - M.Sc. in Biochemistry at University College London
1962-1965 - Ph.D. in Plant Biochemistry at Jesus College, Cambridge
1965-1966 - 1 year Overseas Research Fellowship (Ministry of Overseas Development) at Tea Research Station, Mlanje (now Mulanje), Malawi
1967-1970 - 3 years within International Biological Programme (IBP) with the U.K. Nature Conservancy working on Productivity of Native Ecosystems, at Moor House Field Station, Cumberland
1971-2001 - U.K. Forestry Commission working on Biochemical and Molecular analysis of native and introduced forest trees.